Monday, September 4, 2017

FISHERY RESOURCES OF INDIA TELUGU



ప్రశ్న వివిధరకాల మత్స్యవనరులను/ఫిషరీ రిసోర్సెస్
గురించి వ్యాసం వ్రాయుము





జ. చేపలు, రొయ్యలు, పీతలు వంటివి జలచర జీవులు.  ఇవి జలావాసాలైన సముద్రం, నదులు, కాలువలు,
చెరువులు, జలాసయాలు, ఉప్పునీటి కయ్యలలో నివసిస్తాయి.  ఈ జీవులు నివసించే ఆవాసాల ఆధారంగా
మత్స్యవనరులను – ఎ. మంచినీటి మత్స్యవనరులు  
బి. బ్రాకిష్ వాటర్/ఉప్పునీటిమత్స్యవనరులు మరియు సి. సాగరజల మత్స్యవనరులు
గా విభజించవచ్చును





ఎ. భారతదేశపు మంచినీటి/ఇన్ లాండ్ మత్స్యవనరులు





బారతదేశపు మొత్తం
మత్స్య ఉత్పత్తులలో 30% మంచినీటి వనరులనుండే వస్తున్నాయి.  భారతదేశపు మంచినీటి వనరులులో 195,210
కిలోమీటర్ల పొడవున నదులు, కాలువలు,  2.9
మిలియన్ హెక్టార్ల విస్తీర్ణంతో జలాసయాలు, 2.4 మిలియన్ హెక్టార్ల విస్తీర్ణంతో
చెరువులు, కుంటలు కలవు.





ప్రధాన మంచినీటి వనరులు
– ఉత్తరభారతదేశములో గంగానది, బ్రహ్మపుత్ర నదులు- దక్షిణభారతదేశములో మహానది,
కృష్ణానది, గోదవరి, కావేరి, నర్మదా, తపతి నదులు మొదలైనవి





గంగానది:


గంగానది హిమాలయాలలో
పుట్టి 12500 కి.మీ లతో,  97.6 మిలియన్ల
హెక్టార్ల విస్తీర్ణంతో భారతదేశంలో ప్రవహించే ఒక జీవనది.  దీనిలో- లేబియో రోహితా, కట్లా కట్లా, సిర్రినస్
మ్రిగిలా, హిల్సా ఇలైషా, వాలగో అట్టు, నోటోప్టెరిస్ చిటాలా వంటి చేపలు ప్రధాన
వనరులు


బ్రహ్మ పుత్ర నది


ఇది 4023 కిమీ పొడవుతో,
51 మిలియన్ హెక్టార్ల విస్తీర్ణంతో ఉత్తరభారతదేశంలో ప్రవహిస్తున్నది.  ఈ నదిలో మత్శ్య సంపద పుష్కలంగా
లభిస్తుంది.  వాలగో అట్టు, లేబియో రోహితా,
మిస్టిస్ రిటా, పుంటియస్ సరానా, నోటోప్టెరిస్ చిటాలా, సిర్రినస్ చిటాలా మొదలైన
చేపలు ముఖ్యమైనవి





నర్మదా నది: ఇది 1280
కిమీ పొడవుతో, మధ్య ప్రదేష్, గుజరాత్ లలో ప్రవహిస్తుంది.  9.4 మిలియన్ హెక్టార్ల విస్తీర్ణంతో ఉంటుంది. 


లేబియో ఫింబ్రియేటస్, L.calabasu,
L.bata,Cirrhinus reba, Puntius sarana
etc, cat fish groups
such as Mystus senghala, M. cavasius, Wallago attu,
Clupisomagarua, other
fish groups like Tor tor; Channaspp, Mastacembalus spp; Notopterus notopterus etc.
వంటి వివిధ చేపలు లభిస్తాయి. 





కావేరి
నది.
  ఇది 800 కిమీ పొడవుతో, 4.7 మిలియన్
హెక్టార్ల విస్
తీర్ణంతో  ఉంది
. The fishes like Tor.
Putitora, Barbus
  dubius, Labeo
kontius,Cirrhinus cirhosa, Mystus seenghala, Pangasius pangasius,Wallago attu
, carps such
as Catla catla; Labeo rohita; Cirrhinus mrigala and the
exoticspecies Cyprinus carpio and Osphron
emus goramy
&  game fish like Tor
khudri
and Tmussullahare  
వంటి చేపలు ఈ నదిలో కనిపిస్తాయి.


తపతి నది:  ఇది 720 కిమీ పొడవుతో ప్రవహిస్తుంది.  ఈ నదిలో ప్రధానంగా Tor tor, Mystusseenghala, Wallago attu, Labeo calabasu,
Labeo fimbriata, Cirrhinus mrigala, Chann
a
spp etc
వంటి
చేపలు లభీస్తాయి.


ఆంధ్రప్రదేష్ లో
ఉన్న నదులు


గోదావరి
నది.   దీనిపొడవు 1465 కి.మీ.  గోదావరి నది కాచ్ మెంట్ ఏరియా 315980
చ.కిమీ.  ఈ నదిలో లభించు చేపలు..
Labeo rohita, L.
calabasu, L. fimbriatus, Catla catla, Cirhinus mirigala, Mystes singhaal, Wallago
attu, Hilsa ilaisha, Bangarius bagarius, 
Macrobrachium rosenbergii etc.
వర్షాకాలంలో హిల్సా చేపలు మంచి ఆదాయవనరుగా
ఉంటాయి.
 


కృష్ణా నది: ఇది 1401 కిమీ పొడవుతో 233229
చకీ. కాచ్ మెంట్ విస్తీర్ణంతో ఉంటుంది.  కృష్ణా
నదిలో లభించు చేపలు గోదావరి మత్స్య సంపద వలెనె ఉంటుంది. 


పెన్నా నది: 
ఇది 600 కిమీ పొడవు ఉంటుంది. 
కార్ప్ చేపలు, పిల్లి చేపలు విరివిగా లభిస్తాయి.





ఉప్పునీటి మత్స్య వనరులు





నది
వెళ్ళి సముద్రంతో కలిసే ప్రాంతాన్ని ఎస్చువరీ లేదా బ్రాకిష్ లేదా ఉప్పునీటి జలవనరులు
అంటారు.  ఈ ప్రాంతములో లవణ సాంద్రత విపరీతమైన
వ్యత్యాసాలను ప్రదర్శిస్తుంది.  సాధారణంగా
ఇది వాతావరణ పరిస్థితులను బట్టి, సముద్రపు పోటును అనుసరించి 5-30 పి.పి.టి మధ్య
ఉంటుంది.  ఏ జీవులైతే లవణ సాంద్రతా
వ్యత్యాసాలను తట్టుకొని బ్రతకగలవో అవే ఇలాంటి జలాలలో జీవించగలవు.  అలాంటి జీవులను యూరీ హేలైన్ జీవులు
అంటారు.  బ్రాకిష్ జలాలు అధిక
పోషకపదార్ధాలను కలిగిఉంటాయి.


భారత
దేశపు ముఖ్యమైన ఉప్పునీటి వనరులు 


ఎ.
హుగ్లి ఎస్చువరీ: ఇక్కడ గంగానది బంగాళాఖాతములో కలుస్తుంది.


బి.
గుజరాత్ లోని మహానది ఎస్చువరీ


సి.
ఆంధ్రప్రదేష్ లోని క్రిష్ణా గోదావరి ఎస్చువరీ


డి.
తమిలనాడులోని కావేరీ ఎస్చువరీ


ఇ.
ఒరిస్సాలోని చిలకా సరస్సు, తమిలనాడులోణి పులికాట్ సరస్సు, ఆంధ్రప్రదేష్ లోని
కొల్లేరు సరస్సులు వంటివి ప్రముఖమైన ఉప్పునీటి సరస్సులు.





ఉప్పునీటి
జలాల లక్షణాలు


ఎ.
ఇక్కడ లభించే జలాలు ఉప్పు మరియు మంచినీళ్ళ మిశ్రమము


బి.
పి.హెచ్.  7.5-8.5 మధ్యలో ఉంటుంది


సి.
సేంద్రియ పదార్ధాలు అధికంగా ఉంటాయి.


ఇ.
ఆక్సిజన్ శాతం తక్కువగా ఉంటుంది.


డి.
ఎస్చువరీలు అనేక చేపలకు ప్రత్యుత్పత్తి స్థలాలుగా ఉంటాయి


ఎఫ్.
ఎస్చువరీలు అనేక అనడ్రామస్, కటాడ్రామస్ చేపలకు నివాసం గా ఉంటుంది.





భారతదేశములో
చేపలపెంపకానికి అనువుగా ఉండే సుమారు 1.2 మిలియన్ హెక్టార్ల బ్రాకిష్ వాటర్ జలాలు
అందుబాటులో ఉన్నాయి.  వీటిలో సుమారు 13%
మాత్రమే వినియోగంలో ఉంది.





ఎస్చువరీలలో  లభించే మత్స్యసంపద


ఎ. ముల్లెట్ లు:  బ్రాకిష్ వాటర్ జలాలో పట్టుకొన్న మత్స్యసంపదలో
సుమారు మూడవవంతు ముల్లెట్లే.  వీటిలో ఈ
క్రింది జాతులు ముఖ్యమైనవి. 
Mugil cephalus,  M. tade, M. cunnesius, Valamugil seheli, Liza
macrolepis, L. tade, L. parsia. etc. .
ఈ చేపలు సంవత్సరం పొడవునా లభిస్తాయి.  క్రిష్ణా ఎస్చువరీలో ముల్లెట్లు ప్రముఖ పాత్ర
వహిస్తాయి.


బి. పెర్చెస్:  లేటిస్ కాల్కెరిఫెర్ ప్రముఖమైన పెర్చ్.  ఇంకా Holocantrus serranus, Lethirinus nebulosus,
Ambassis ambassis, Terapon jarbua  Etc.


సి. పిల్లి చేపలు:  ఇవి చాలా ముఖ్యమైన చేపలు.  వీటిని బ్రాకిష్ జలాలలో పెంపకం కూడా
చేస్తున్నారు.  ముఖ్యమైన జాతులు
Mystus gulio,
Heteropneustes fossilis, Pangasius sutchi, P. pangasius Arieus, Myceous sp.
etc.


డి. క్లుపియాయిడ్ లు:  వీటిలో ప్రముఖమైనది హిల్సా ఇలైషా.  మంచి మార్కెట్ గిరాకీ కలిగిన చేప.  హుగ్లీ ఎస్చువరీలో హిల్సా ప్రముఖమైన దిగుబడి.  ఇది ఒక అనాడ్రొమస్ చేప.  Neotilosa, Manon, Elops sp. వంటివి ఇతర
క్లుపియాయిడ్ లు.





ఎస్చువరీలలో లభించే రొయ్యలు పీతలు


ఎస్చువరీలలో లభ్యమయ్యే షెల్ ఫిషరీ (రొయ్యలు,
పీతలు) రొయ్యలలో  ముఖ్యమైనవి


Pinaeus indicus, 
P. monodon (Tiger prawn), P. semiselcatus, Metapenaeus dobsonii, M.
monoceros, M. brevicornis
మొదలగునవి. పీతలలో ముఖ్యమైనవి  Scyla serrata, Portunus pelagicus వంటివి.





Saturday, September 2, 2017

PLANKTON PRACTICAL MATERIAL

1. Coscinodiscus
Classification:
Empire Eukaryota
Kingdom Chromista
Phylum Bacillariophyta
Subphylum Coscinodiscophytina
Class Coscinodiscophyceae
Subclass Coscinodiscophycidae
Order Coscinodiscales
Family Coscinodiscaceae
Genus Coscinodiscus
1. This is centric diatom
2. Cells are discoid and wedge-shaped, visible in girdle view with one side
higher than the other (may not be visible in all orientations).
3. Valves are most convex at the tallest part of the cell, near one end of the cell. Cells have a rosette of larger areolae at the centre of the valve.
4. Chloroplasts are smooth and disc-shaped.
5. Cells are yellow-brown in colour."About 8 areolae in 10 μm near centre, 10 midway to margin, and 11 near margin; on edge of valve mantle 13 in 10 μm.
6. Chamber openings small, dotlike. Outer closing membrane of areolae very
delicately poroid. Radial rows and secondary spiral rows distinct. Marginal spinulae and the hyaline lines radiating from the spinulae toward the center distinct, 5 - 7 μm apart. Two small processes or apiculi on margin at distance of about 120° from each other. Girdle formed from the two similar. No intercalary bands".

2. Chaetoceros
Empire Eukaryota
Kingdom Chromista
Subkingdom Harosa
Infrakingdom Heterokonta
Phylum Ochrophyta
Subphylum Khakista
Class Bacillariophyceae
Subclass Coscinodiscophycidae
Order Chaetocerotales
Family Chaetocerotaceae
Genus Chaetoceros
1. Cells are usually connected in chains, and are roughly rectangular in
girdle view and elliptical in valve view (elliptic cylinder),
2.Two spines arising from each valve.
3.Adjacent cells are linked by the crossing or touching of the spines near the base.
4.Cells are yellow-brown in colour.


3. Biddulphia
Classification:
Empire Eukaryota
Kingdom Chromista
Phylum Bacillariophyta
Subphylum Bacillariophytina
Class Mediophyceae
Subclass Biddulphiophycidae
Order Biddulphiales
Family Biddulphiaceae
1. Cells robust, rectangular in girdle view, elliptical in valve view, with prominent elevations at the poles.
2. Normally seen in girdle view, often growing in zig-zag chains attached to filamentous seaweeds, etc. Often found also in inshore plankton samples. Plastids numerous, discoidal.
3. A very common marine genus but taxonomically extremely confused (see below). Valves bipolar/lanceolate to almost circular, often with waxy margins. Valves surface often furrowed with various thickenings, spines (often conspicuous in the central region) or ridges.
4. Apices bearing rounded pseudocelli on low or extended elevations. Valve mantle not well-defined but extreme edge of valve often recurved and variously moulded. Areolae large with perforate vela of the cribrum type. Simple pores also occur occasionally in the valve framework.
5. Internally with conspicuous plain ridge (pseudosepta) beneath the external indentations of the valve.
6. One to several rimoportulae present, sessile, clustered in centre; external tubes often stout and surmounted by two spines. Cingula with a complete, close valvocopula and 3-4 split copulae; areolae large, in rows; fluting of girdle corresponding to that of valve edge.

4. Skeletonema

Phylum Bacillariophyta
Subphylum
Class Coscinodiscophyceae
Subclass Thalassiosirophycidae
Order Thalassiosirales
Family Skeletonemaceae
Genus Skeletonema

1.Cells are short and cylindrical, usually connected in long, straight or slightly undulate chains by a marginal ring of spines (strutted processes).
2.The valve face is convex to flat. Spines interlock midway between adjacent cells, visible as a dotted ring; spine lengths are variable (Horner.
3.There are two chloroplasts per cell and the nucleus is located centrally. Strutted processes are tubular but semicircular in cross section.

4.One labiate process is present near the center of the valve inside the ring of processes.
5.Molecular data has recently separated S. costatum into several species. These distinctions cannot be made without detailed examination of their morphology.

5. Leptocylindrus


Phylum Ochrophyta
Subphylum Khakista
Class Coscinodiscophyceae
Subclass Chaetocerotophycidae
Order Leptocylindrales
Family Leptocylindraceae
Genus Leptocylindrus
1.Cells are cylindrical and form long, straight chains. Cells are connected by the whole valve surface.
2.The central parts of the valve face may be slightly convex/concave, fitting into the concavity/convexity of an adjacent valve.
3.Cells are thin-walled and do not have any obvious spines or processes. Using SEM, short flap-like spines are visible on the border between the valve face and mantle (Hasle and Syvertsen 1997).
4.Chloroplasts are numerous small ovoid plates, distributed throughout the cell (Cupp 1943). Intercalary bands are not visible with LM.
5. Resting spores form as a result of sexual reproduction, they consist of two unequal valves and are found in an auxospore.
6.Resting spores are covered with spicules.

6. Pleurosigma
Phylum Bacillariophyta
Subphylum Bacillariophytina
Class Bacillariophyceae
Subclass Bacillariophycidae
Order Naviculales
Suborder Naviculineae
Family Pleurosigmataceae
Genus Pleurosigma
1. Valves are elongate and sigmoid. Striae are composed of evenly spaced decussate rows.
2. The rows are arranged in 3 patterns: a trans-apical row and two opposing oblique rows.
3. Internally, the areolae are occluded by hymenes. The raphe is strongly sigmoid, with a narrow axial area.
4. The sternum is thickened equally on both sides. The raphe terminates proximally in a small oval, expanded central area defined by two curved ridges.
5. Proximal raphe ends are slightly inflated and straight, or unilaterally bent. The distal raphe ends are hooked and deflected to opposite sides on the mantle.
6. The valve mantle is relatively shallow and the valve / mantle interface is broad and curved. Two to four ribbon-like plastids extend from apex to apex.



7. Asterionella

Phylum Ochrophyta
Subphylum Khakista
Class Bacillariophyceae
Subclass Fragilariophycidae
Order Fragilariales
Family Fragilariaceae
Genus Asterionellopsis

1.Cells are needle-shaped and arranged in star-like and helical chains. In girdle view, cells have a narrow and elongated neck with a broad triangular base.
2. The end of the neck is not lobed. Adjacent cells are attached by the valve face of the base; therefore cells are usually seen in girdle view.
3. In valve view, the base is lobed. One or two plate-like chloroplasts are located in the base (Cupp 1943). Cells are yellow-brown in colour.
4. Characteristics observable under SEM: "Slit fields or apical pores at both ends of valve.
5. A single labiate process present at the narrow end of the valve. Transapical striae are very delicate.


8. Thalassionema
Classification:
Empire Eukaryota
Kingdom Chromista
Phylum Bacillariophyta
Subphylum Bacillariophytina
Class Bacillariophyceae
Subclass Fragilariophycidae
Order Thalassionematales
Family Thalassionemataceae
Genus Thalassionema
1.Cells are usually in star-shaped or zigzagged chains connected by mucilage pads on the ends of valves. Cells are rectangular in girdle view, with valve ends that are similar in shape and width, and are narrowly elliptical in valve view.
2.Chloroplasts are small and numerous.
3.Cells are yellow-brown in colour (Guiry 2011).Each valve has two labiate processes, one on each end; a small apical spine is sometimes also present.
3."Marginal areolae are visible as ribs with LM.
4.The sternum is broad and without areolae.
5.Internally, each valve pole has a simple, slit-like labiate process, which appears externally as a small, round hole.

9. Thalassiothrix


Classification:
Empire Eukaryota
Kingdom Chromista
Phylum Bacillariophyta
Subphylum Bacillariophytina
Class Bacillariophyceae
Subclass Fragilariophycidae
Order Thalassionematales
Family Thalassionemataceae
Genus Thalassiothrix

1. Cells needle-like, straight, slightly curved or sigmoidal in girdle view, forming radiating colonies, joined by bent foot poles (one end of frustule); head poles separated.
2. Apical axis 420–5680 µm; transapical axis 1.5–6 µm; foot pole apexrounded; head pole apex rounded but with 2 protruding spines.
3. Areolae in a single marginal row, 12–17 in 10 µm, with complex vela. Row of fine spines, marginal on the vela, pointed towards the head pole, 1–2 in 10 µm, occurring along the frustule but more abundant towards the head pole. Small foramina on valve surface.
4. Labiate process at each apex.
5. Chloroplasts numerous, spherical.


10. Amphora

Phylum Bacillariophyta
Subphylum Bacillariophytina
Class Bacillariophyceae
Subclass Bacillariophycidae
Order Thalassiophysales
Family Catenulaceae
Genus Amphora

Solitary cells that can be motile and almost always occur in girdle view. The cells appear elliptical with flat truncate ends. The cell appears as a large slice of orange. The valves are asymmetrical and are sometimes smaller or constricted at each end of the cell. Both rapheslie on the same side of the valve. There are generally two or more plastids that occur in different positions throughout the cell.

Epiphytic on plants, stones and mud. A large mainly marine genus with relatively few species in freshwater.



11. CERATIUM
Phylum Dinoflagellata
Subphylum
Class Dinophyceae
Subclass
Order Gonyaulacales
Family Ceritiaceae
Genus Ceratium

1.Ceratium furca has a wide girdle and a prominent straight apical horn.
2.C. furca also has two unequal posterior horns.
3.The right horn is shorter than the left.
4.There is a thin bar that connects the two horns.
5.C. furca is yellow-brown and has thick thecal plates.


12. Protoperidinium

Phylum Myzozoa
Subphylum Dinoflagellata
Class Dinophyceae
Subclass
Order Peridiniales
Family Protoperidiniaceae
Genus Protoperidinium

1.Protoperidinium oceanicum cell is star-shaped with one long apical
horn and two long antapical horns.
2.The centre of the cell theca is round with a protrusion on each side.
3.The cell cingulum is narrow and has wide lists.
4.The two antapical horns are long, tubular, pointed and divergent.
5. The left antapical horn is a bit shorter and thinner than the right. Cells have a deep sulcus that forms a strong indentation between the horns.
6.The theca is reticulated with spiny junctions, making it look very ornate under SEM.


13. Dinophysis

Phylum Dinoflagellata
Subphylum
Class Dinophyceae
Subclass
Order Dinophysiales
Family Dinophyciaceae
Genus Dinophysis

1.Cells are broadly subovoid and widest posteriorly.
2.Cells have a curved dorsal margin and an almost straight ventral margin. The cell surface has deep poroids.
3.Cells also have two well-developed sulcal lists.
4.The left sulcal list is about ⅘ of the cell length.
5.The right sulcal list is also long and can extend beyond the second rib.
6.Dinophysis fortii has a wide rounded posterior and reticulation on the sulcal lists. D. fortii cells have many large centrally placed chloroplasts.



Trichodesmium
Kingdom: Bacteria

Phylum: Cyanobacteria

Order: Oscillatoriales

Genus: Trichodesmium

Trichodesmium, also called sea sawdust, is a genus of filamentous cyanobacteria. They are found in nutrient poor tropicaland subtropical ocean waters (particularly around Australia and in the Red Sea, where they were first described by Captain Cook). Trichodesmium is a diazotroph; that is, it fixes atmospheric nitrogen into ammonium, a nutrient used by other organisms.Trichodesmium is the only known diazotroph able to fix nitrogen in daylight under aerobic conditions without the use ofheterocysts.[1]
Trichodesmium can live in solitude or in colonies. These colonies are visible to the naked eye and sometimes form blooms, which can be extensive on surface waters. These large blooms led to widespread recognition as "sea sawdust/straw"; in fact, the Red Sea gets most of its eponymous colouration from the corresponding pigment in Trichodesmium erythraeum. Colonies of Trichodesmium provide a pseudobenthic substrate for many small oceanic organisms including bacteria, diatoms,dinoflagellates, protozoa, and copepods (which are its primary predator); in this way, the genus can support complex microenvironments.
Like most cyanobacteria, Trichodesmium has a gram negative cell wall. However, unlike other aerobic diazotrophs, heterocysts (structures found in cyanobacteria which protect nitrogenase from oxygenation) are lacking in Trichodesmium. This is a unique characteristic among aerobic diazotrophs which fix nitrogen in daylight. Photosynthesis occurs using phycoerythrinlight harvesting phycobiliprotein which is normally found within heterocysts in other diazotrophs.
Instead of having localized stacks of thylakoids, Trichodesmium has unstacked thylakoids found throughout the cell. Trichodesmium is highly vacuolated and the content and size of the vacuoles shows diurnal variation. Large gas vesicles (either along the periphery as seen in T. erythaeum or found distributed throughout the cell as seen in T. thiebautii) allow Trichodesmium to regulate buoyancy in the water column. These gas vesicles can withstand high pressure, presumably those up to 100 – 200 m in the water column, allowing Trichodesmium to move vertically through the water column harvesting nutrients.

Spirulina
Domain: Bacteria

Kingdom: Eubacteria

Phylum: Cyanobacteria

Order: Spirulinales

Family: Spirulinaceae

Genus: Spirulina






Spirulina is a genus of blue-green algae used as a nutritional supplement. Blue-green algae, which are microscopic fresh-water organisms, are also known as cyanobacteria. Their color is derived from the green pigment of chlorophyll, and the blue from a protein called phycocyanin. The species most commonly recommended for use as a nutritional supplement are Spirulina maxima and Spirulina platensis. These occur naturally in warm, alkaline, salty, brackish lakes, but are also commonly grown by aquaculture and harvested for commercial use. Spirulina contains many nutrients, including B vitamins, beta-carotene, gamma-linolenic acid, iron, calcium, magnesium, manganese, potassium, selenium, zinc, bioflavonoids , and protein.
Spirulina is about 65% protein by composition. These proteins are complete, in that they contain all essential amino acids , plus some nonessential ones. In that regard, it is similar to animal protein, but does not contain saturated fats, or residues of hormones or antibiotics that are in some meats. Since spirulina is normally taken in small amounts, the quantity of dietary protein supplied for the average reasonably well-nourished person would not be significant. However, it is a good source of trace minerals, some vitamins, bioflavonoids, and other phytochemicals. It also has high digestibility and bioavailability of nutrients.


Nostoc sp
Kingdom: Bacteria

Phylum: Cyanobacteria

Class: see taxonomic note

Order: Nostocales

Family: Nostocaceae

Genus: Nostoc


Nostoc is a genus of cyanobacteria found in a variety of environmental niches that forms colonies composed of filaments of moniliform cells in a gelatinous sheath.
The name Nostoc was coined by Paracelsus.[1]
Nostoc can be found in soil, on moist rocks, at the bottom of lakes and springs (both fresh- and saltwater), and rarely in marine habitats. It may also grow symbiotically within the tissues of plants, such as the evolutionarily ancient angiospermGunnera and the hornworts (a group of bryophytes), providing nitrogen to its host through the action of terminally differentiated cells known as heterocysts. These bacteria contain photosynthetic pigments in their cytoplasm to perform

Anabaena
Kingdom: Bacteria

Phylum: Cyanobacteria

Order: Nostocales

Family: Nostocaceae

Genus: Anabaena
Anabaena is a genus of filamentous cyanobacteria that exists as plankton. It is known for its nitrogen fixing abilities, and they form symbiotic relationships with certain plants, such as the mosquito fern. They are one of four genera of cyanobacteria that produce neurotoxins, which are harmful to local wildlife, as well as farm animals and pets. Production of these neurotoxins is assumed to be an input into its symbiotic relationships, protecting the plant from grazing pressure.
A DNA sequencing project was undertaken in 1999, which mapped the complete genome of Anabaena, which is 7.2 million base pairs long. The study focused on heterocysts, which convert nitrogen into ammonia. Certain species of Anabaena have been used on rice paddy fields, proving to be an effective natural fertilizer.


Copepoda
Kingdom: Animalia

Phylum: Arthropoda

Subphylum: Crustacea

Class: Maxillopoda

Subclass: Copepoda

Copepods vary considerably, but can typically be 1 to 2 mm (0.04 to 0.08 in) long, with a teardrop-shaped body and large antennae. Although like other crustaceans, they have an armoured exoskeleton, they are so small that in most species, this thin armour, and the entire body, is almost totally transparent. Some polar copepods reach 1 cm (0.39 in). Most copepods have a single median compound eye, usually bright red and in the centre of the transparent head; subterranean species may be eyeless. Like other crustaceans, copepods possess two pairs of antennae; the first pair is often long and conspicuous.
Copepods typically have a short, cylindrical body, with a rounded or beaked head. The head is fused with the first one or two thoracic segments, while the remainder of the thorax has three to five segments, each with limbs. The first pair of thoracic appendages is modified to form maxillipeds, which assist in feeding. The abdomenis typically narrower than the thorax, and contains five segments without any appendages, except for some tail-like "rami" at the tip.[4]
Because of their small size, copepods have no need of any heart or circulatory system (the members of the order Calanoida have a heart, but no blood vessels), and most also lack gills. Instead, they absorb oxygen directly into their bodies. Their excretory system consists of maxillary glands.


Amphipods
Kingdom: Animalia

Phylum: Arthropoda

Subphylum: Crustacea

Class: Malacostraca

Superorder: Peracarida

Order: Amphipoda



The body of an amphipod is divided into 13 segments, which can be grouped into a head, a thorax and an abdomen.[4]
The head is fused to the thorax, and bears two pairs of antennae and one pair of sessilecompound eyes.[6] It also carries the mouthparts, but these are mostly concealed.[7]
The thorax and abdomen are usually quite distinct and bear different kinds of legs; they are typically laterally compressed, and there is no carapace.[6] The thorax bears eight pairs of uniramous appendages, the first of which are used as accessorymouthparts; the next four pairs are directed forwards, and the last three pairs are directed backwards.[6] Gills are present on the thoracic segments, and there is an open circulatory system with a heart, using haemocyanin to carry oxygen in the haemolymphto the tissues. The uptake and excretion of salts is controlled by special glands on the antennae.[4]
The abdomen is divided into two parts: the pleosome which bears swimming legs; and the urosome, which comprises a telsonand three pairs of uropods which do not form a tail fan as they do in animals such as true shrimp.





Lucifer
Phylum Arthropoda
Subphylum Crustacea
Class Malacostraca
Superorder Eucarida
Order Decapoda
Suborder Dendrobranchiata
Superfamily Sergestoidea
Family Luciferidae
Genus Lucifer














 Sometimes called the ghost shrimp.
 Appearance of an elongated shrimp; it almost seems to have been stretched.
 Carapace is extremely laterally compressed.
 Head section is longer and narrower than the thorax, resulting in the eyes and antennae being widely distant from the mouthparts.
 Rostrum is short and pointed.
 3 pairs of pereiopods instead of the usual 5; the 3rd pair is partially chelate .
 Telson in males has 2 distinct projections on the ventral surface.
 Almost transparent.
Distribution
 South-eastern Australian waters (Dakin and Colefax 1940). Melbourne Harbour (Borradaille 1916). South-eastern Tasmanian waters and Derwent Estuary (Ong 1967, Nyan Taw 1978). Indian Ocean and South China Sea (Hansen 1919).
 Particularly abundant in the coastal waters of south-eastern Australia where it can often make sorting of other plankton difficult because of the large numbers.
 Nyan Taw (1978) reported that it is a dominant species in the inshore coastal waters of south-eastern Tasmania during winter months.

Euphasiid

Kingdom: Animalia

Phylum: Arthropoda

Subphylum: Crustacea

Class: Malacostraca

Superorder: Eucarida

Order: Euphausiace
. The profuse exposed gills of euphausiids arise from the coxae of the thoracic legs (feathery entirely exposed). They are conspicuous and are progressively larger toward the posterior of the thorax. In mysids and in the penaeid and caridean decapods the gills, when present, arise beneath the carapace, are relatively compact, and only partly exposed (mostly hidden beneath carapace).

2. In euphausiids and decapods, the carapace is firmly attached to all eight thoracic segments. In mysids, the carapace is attached only to the three anterior segments of the thorax.

3. The rostrum and frontal plate in euphausiids and mysids are parts of a basically horizontal projection of the carapace. In the decapods (penaeids and carideans) the frontal plate supports an elevated, laterally compressed rostrum, often acute and anteriorly or dorsally notched.

4. The antennae. The 3-segmented peduncle of the 1st antenna in euphausiids, particularly the lappet on segment 1, provides important taxonomic characters. In pelagic decapoods and mysids the peduncles are usually relatively simple and uniform. In the larger nektonic species of sergestid decapods, the flagella of the 1st antenna can be extremely elongate and thread-like, and the flagellum of the 2nd antenna can be uniquely coiled like a spring.

5. The abdominal pleura. These are lateral coverings of abdominal segments 1-5 in euphausiids and pelagic decapods. They consist of plates, with free ventral edges (pleuron seg. 2 overlaps seg. 3). In most mysids the abdominal segments are nearly cylindrical, without lateral plates. Caridean decapods are unique in that the pleura of the 2nd abdominal segment overlap pleura of both the 1st and 3rd segments (pleuron seg 2 overlaps seg 1&3).

6. Statocysts. The broad proximal ends of the endopods of the uropods in mysids (except the predominantly deep living Lophogastridae and one small family of Mysida) have conspicuous circular statocysts (uropods with statocyst). Sergestid decapods have statocysts, but they are at the bases of the peduncles of the 1st antennae . Statocysts are not known in euphausiids (uropods without statocyst).

7. The telson of adult euphausiids is about as long as the exopod of the uropod, and has an acute distal point which is flanked by a pair of very strong posterior lateral spines which extend beyond the telson tip. In mysids the telson is shorter than the uropods, paddle-like and spinose in some species, rarely with an acute tip, and in many genera it is cleft distally, appearing as a two-pronged, sometimes spinose or setose fork. In sergestid decapods the tip of the telson is bluntly rounded, shorter than the uropod, and marginally setose.


Mysid

Kingdom: Animalia

Phylum: Arthropoda

Subphylum: Crustacea

Class: Malacostraca

Subclass: Eumalacostraca

Superorder: Peracarida



The head of a mysid bears two pairs of antennae and a pair of large, stalked eyes. The head and first segment (or sometimes the first three segments) of the thorax are fused to form the cephalothorax. The eight thoracic segments are covered by the carapace which is attached only to the first three. The first two thoracic segments bearmaxillipeds which are used to filter plankton and organic particulate from the water. The other six pairs of thoracic appendages are biramous (branching) limbs known aspereopods, and are used for swimming, as well as for wafting water towards the maxillipeds for feeding. Unlike true shrimps (Caridea), females have a marsupiumbeneath the thorax. This brood pouch is enclosed by the large, flexible oostegites, bristly flaps which extend from the basal segments of the pereopods and which form the floor of a chamber roofed by the animal's sternum. This chamber is where the eggs are brooded, development being direct in most cases.[2]
The abdomen has six segments, the first five of which bear pleopods, although these may be absent or vestigial in females. The fourth pleopod is longer than the others in males and has a specialized reproductory function.[2]
The majority of species are 5–25 mm (0.2–1.0 in) long, and vary in colour from pale and transparent, through to bright orange or brown. They differ from other species within the superorder Peracarida by featuring statocysts on their uropods (located on the last abdominal segment). These help the animal orient itself in the water and are clearly seen as circular vesicles: together with the pouch the statocysts are often used as features that distinguish mysids from other shrimp-like organisms.[3]


Pteropod

Kingdom: Animalia

Phylum: Mollusca

Class: Gastropoda

Clade: Heterobranchia

Informal group: Opisthobranchia

Informal group: Pteropoda
Pteropoda (common name pteropods, from the Greek meaning "wing-foot") are specialized free-swimming pelagic sea snails and sea slugs, marine opisthobranch gastropods. The monophyly of Pteropoda is the subject of a lengthy debate; they have even been considered as paraphyletic with respect to cephalopods.[1] Current consensus, guided by molecular studies, leans towards interpreting the group as monophyletic.[2]
Pteropoda encompasses the two clades Thecosomata, the sea butterflies, and Gymnosomata, the sea angels. The Thecosomata (lit."case-body"[3]) have a shell, while the Gymnosomata ("naked body") do not. The two clades may or may not be sister taxa; if not, their similarity (in that they are both pelagic, small, and transparent, and both groups swim using wing-like flaps (parapodia) which protrude from their bodies) may reflect adaptation to their particular lifestyle.


Ostracoda

Kingdom: Animalia

Phylum: Arthropoda

Subphylum: Crustacea

Class: Ostracoda










The body of an ostracod is encased by two valves, superficially resembling the shell of a clam. A distinction is made between the valve (hard parts) and the body with its appendages (soft parts).
Soft parts[edit]
The body consists of a head and thorax, separated by a slight constriction. Unlike many other crustaceans, the body is not clearly divided into segments. The abdomen is regressed or absent, whereas the adult gonads are relatively large.
The head is the largest part of the body, and bears most of the appendages. Two pairs of well-developed antennae are used to swim through the water. In addition, there is a pair of mandibles and two pairs of maxillae. The thorax typically has two pairs of appendages, but these are reduced to a single pair, or entirely absent, in many species. The two "rami", or projections, from the tip of the tail, point downwards and slightly forward from the rear of the shell.[14]
Ostracods typically have no gills, instead taking in oxygen through branchial plates on the body surface. Most ostracods have no heartor circulatory system, and blood simply circulates between the valves of the shell. Nitrogenous waste is excreted through glands on the maxillae, antennae, or both.[14]
The primary sense of ostracods is likely touch, as they have several sensitive hairs on their bodies and appendages. However, they do possess a single naupliar eye, and, in some cases, a pair of compound eyes, as well.[14]


Cladocera
Kingdom: Animalia

Phylum: Arthropoda

Subphylum: Crustacea

Class: Branchiopoda

Subclass: Phyllopoda

Order: Cladocera














They are mostly 0.2–6.0 mm (0.01–0.24 in) long, with the exception of Leptodora, which can be up to 18 mm (0.71 in) long.[1] The body is not obviously segmented and bears a folded carapace which covers the thorax and abdomen.[2]
The head is angled downwards, and may be separated from the rest of the body by a "cervical sinus" or notch.[2] It bears a single black compound eye, located on the animal's midline, in all but two genera, and often, a single ocellus is present.[3] The head also bears two pairs of antennae – the first antennae are small, unsegmented appendages, while the second antennae are large, segmented, and branched, with powerful muscles.[2] The first antennae bear olfactory setae, while the second are used for swimming by most species.[3] The pattern of setae on the second antennae is useful for identification.[2] The part of the head which projects in front of the first antennae is known as the rostrum or "beak".[2]
The mouthparts are small, and consist of an unpaired labrum, a pair of mandibles, a pair of maxillae, and an unpaired labium.[2]They are used to eat "organic detritus of all kinds" and bacteria.[2]
The thorax bears five or six pairs of lobed, leaf-like appendages, each with numerous hairs or setae.[2] Carbon dioxide is lost, and oxygen taken up, through the body surface.[2]

Friday, September 1, 2017

CRUSTACEAN LARVAE

• 1 CRUSTACEAN LARVAE
o 1.1 INTRODUCTION
 1.1.1 NAUPLIUS
 1.1.2 METANAUPLIUS
 1.1.3 CYPRIS
 1.1.4 'PROTOZOEA
 1.1.5 ZOEA
 1.1.6 METAZOAEA
 1.1.7 CALYPTOSIS
 1.1.8 ERICHTHUS
 1.1.9 ALIMA
 1.1.10 MEGALOPA
 1.1.11 GLAUCOTHOEA
 1.1.12 MYSIS
 1.1.13 PHYLLOSOMA
CRUSTACEAN LARVAE
INTRODUCTION
Crustaceans, are large number of arthropods, comprising almost 52,000 described species. The majority of them are aquatic living in either marine or fresh water environments, but a few groups have adapted to life on land, such as terrestrial crabs , terrestrial hermit crabs and wood lice. Crustaceans show both direct and indirect development. In most crustacea, development is accompanied by little or more metamorphosis and the various stages of development are known as larvae. Several larval forms are met within Crustacea and special terms are applied to each one of them
NAUPLIUS


It is the simplest, commonest and earliest larval form in crustacea. Nauplius is a microscopic, oval or pear shaped with an unsegmented body having a broad anterior head region, an intermediate trunk-region and a posterior bilobed anal region. It has 3 pair of unjointed appendages bearing swimming seate. The first pair is uniramous and become the antennules of the adult. Second is antennaryand third is mandibular called antennae and mandibles respectively in the adult. The head region bears a conspicuous sessile median eye. The mouths open anteriorly between the bases of antennary and mandibular feet. While the anus lies at the extremity of the caudal region. The alimentary canal is straight and made of foregut, midgut and hindgut. However mouth and alimentary canal are lacking in nauplius of Cirripedia. The larva is without a heart and a segmented ventral nerve cord.

METANAUPLIUS

It follows the nauplius, is a vaguely defined stage. It consists of an oval cephalothorax, an elongated trunk-region and an abdomen terminating in a caudal fork provided with setae. Dorsal shield of the head grows back to form carapace. In addition to the 3-origional appendages of nauplius, it also develops the rudiments of 4 pairs of appendages, which become the maxillulae the maxillae and the first 2 pairs of maxillipedes of the adult.
Branchiopoda, Cephalocardia and perhaps some Stomatopoda hatch the metanaupliu
CYPRIS


In some Cirripedia (Sacculina, Lepas), the nauplius passes into the cypris stage, in which the body and the appendages are enclosed within bivalved carapace with an adductor muscle to close it. Its modified antennules have of cement glands at their bases. It undergoes a remarkable series of metamorphoses to become the sessile adult form. In Ostracoda, the eggs typically hatch in the cypris form.
PROTOZOEA
The metanauplius larva is succeeded by the protozoaea stage with 7 pairs of appendages and the beginning of segmentation. The carapace become enlarged and covers the dorsal surface anteriorly . The 7 pairs of appendages present in the metanauplius (up to 2nd maxillipede) become well-developed and capable of movements .The rudiments of paired lateral eye begin to appear near the median eye. The rudiments of the remaining posterior six thoracic segments are also marked off, but the abdomen is still unsegmented and without limbs. The protozoaea swims by antennae. In Penaeus hatch in the protozoaea.
ZOEA
Zoaea is the second important larvae of the Crustacea, the first being the nauplius. Protozoaea stage is succeeded by the zoaea stage. The zoaea is characterized with a distinct cephalothorax and abdomen, 8 pair of appendages and buds of 6 more, and resembles the adult Cyclops. The cephalothorax is immensely developed and covered by a helmet-like carapace, which is produced into two long spines, an anterior median rostral and a posterior median dorsal. Two lateral spines are usually also present. The paired lateral and stalked compound eyes become well formed and remaining 6 pair of thoracic appendages appears in the form of bud. The long abdomen is distinctly made of 6 segments, and terminates in a caudal furca, but still lacking in appendages. Zoaea swims by means of thoracic limbs.

METAZOAEA
The older zoaea or metazoaea has well formed third maxillipedes, which are biramous and swimming organs in Anomura, but uniramous and non-swimming in Brachyura. The 6 pairs of abdominal appendages also appear in the form of buds.
CALYPTOSIS
In Euphausiacea, one of the larval stages is termed calyptopsis .It is similar in all respects to a typical zoaea except that the paired 'eyes are not stalked but sessile.
ERICHTHUS
In erichthus larva, a carapace covers the greater part of the body. Head is unsegmented, bearing median and paired eyes and all the 5 pairs of cephalic appendages. The thorax is made of segments, free from the carapace ,and bearing anterior 5 pairs of biramous swimming appendages .The broad abdomen is unsegmented and with a single pair of appendages. Such larvae are met with in Lysiosquilla.
ALIMA
The so-called alima larva of Squilla which hatch out from the egg directly, is a modified zoaea It is apeagic larva, having a glass-like transparency and occurring in large numbers in the plankton. It has a slender form, and a sort and broad carapace. All the head appendages are present. But only is 6-segmented, having 4 or 5 pairs of pleopods. The alima larva differs from the zoaea larva in the armature of the telson and a very large raptorial second maxillipedes.

MEGALOPA


In true crabs, the zoaea larva or metazoaea passes through successive moults into the post larval megalopa stage. It has a broad and crablike unsegmented cephalothorax. The carapace is produced anteriorly into a median spine. The eyes are large, stalked and compound. All the thoracic appendages are well formed of which the last 5 pairs are uniramous. The abdomen is also well formed, straight and bears biramous pleopods.
GLAUCOTHOEA
In hermit crabs ,the metazoaea leads to the glaucothoe stage. It corresponds to the megalopa stage of Bracchyura with a large symmetrical abdomen and a full complement of adult appendages.
MYSIS
In Penaeus, the zoaea , instead of converting into the megalopa stage, moults into the postlarval mysis larva with 13 pairs of appendages.all the thoracic appendages are biramous. Even the 5 pairs of posterior thoracic legs are biramous with flagellar exopodites which take up the locomotory function uptill now now fulfilled chiefly by the antennae. The abdomen develops similar to that of the adult form, with 5 pairs of biramous pleopods and a pair of uropods and telson .the mysis larva metamorphosis in to the adult prawn by the loss of the exopodites on the thoracic legs.
PHYLLOSOMA
In the rock- lobster (palinurus), the newly hatched larva, called the phyllosoma or glass- crab, is a greatly modified mysis stage. It is a remarkable for its large size, extremely flattened and leaf- like delicate form and glassy transparency. A narrow constriction demarcates the head from thorax. A large oval carapace covers the head and the first two thoracic segments. The eyes are compound and borne by large stalks. Only anterior 6 pairs of thoracic appendages are present in the newly hatched larva. The first thoracic appendages or maxillipedes are rudimentary (palinurus) or absent (Scyllarus) and the second are uniramous, succeed by 4 pairs of very long and biramous legs with notatory exopodites Last two pairs of thoracic appendages are usually absent. Abdomen, though indistinctly segmented is very small and limbless. Phyllosoma undergoes several moults before reaching the adult form.



















Larval Forms Found in Crustacea | Invertebrate Zoology
The following points highlight the nine important larval forms found in Crustacea. The larval forms are: 1. Nauplius Larva 2. Metanauplius Larva 3. Protozoaea Larva 4. Zoaea Larva 5. Cypris Larva 6. Mysis or Schizopod Larva 7. Megalopa Larva 8. Phyllosoma Larva 9. Alima Larva.
Larval Form
1. Nauplius Larva:
Nauplius larva is egg-shaped and un-segmented. It has a broad anterior end with a median eye, large labrum and three paired appendages.
The median eye is characteristic of the nauplius larva and is often referred to as the nauplius eye, it is made usually of three but at times four ocelli which are pigmented cups with no lens, and are innervated by the protocerebrum. The median eye may degenerate or persist in the adult crustacean.
The appendages are uniramous antennules having two groups of sensory cells forming frontal organs, a pair of biramous antennae, and a pair of biramous mandibles for swimming, they have gnathobases directed towards the mouth, though the gnathobases of mandibles may be absent at first. A stomodaeum with mouth, proctodaeum with anus, and a midgut are also present.
A typical crustacean hatches as a free-swimming nauplius, but in Malacostraca (except in primitive forms) the nauplius is passed over as a stage within the egg membrane.
However, in certain crustaceans like Branchiopoda the nauplius metamorphoses directly into the adult but in majority of crustaceans it metamorphoses to adult through various intermediate larval stages like metanauplius, protozoaea, zoaea, cypris, mysis, megalopa, phyllosoma, alima, etc.
Larval Form # 2. Metanauplius Larva:
Metanauplius larva is like a nauplius, except that it shows some segmentation of the body, and there are four pairs of additional appendages of the thorax which shows some segmentation; these appendages are two pairs of maxillae and two pairs of maxillipedes. Some Notostraca, such as Apus, hatch as a metanauplius larva.

Larval Form # 3. Protozoaea Larva:
In marine prawns like Penaeus and some other decapods, the nauplius directly develops into protozoaea larva. The body of protozoaea is divisible into cephalothorax and abdomen. The cephalothorax is broad, segmented and covered with carapace.
The appendages that appeared in metanauplius become well developed and functional. The rudiments of other thoracic appendages also appear. The abdomen is unsegmented, without any appendage and has a forked telson.
Larval Form # 4. Zoaea Larva:
Zoaea larva has a well formed head with a long, median dorsal spine, two stalked compound eyes and one simple eye, all appendages from antennules to the last pair of maxillipedes are present, carapace is well formed and produced in front into a rostrum.
Thorax is un-segmented and rudimentary at its hinder end. Abdomen is well formed and six segmented, but it has no appendages except a forked telson. It swims by its biramous maxillipedes.
In Penaeus, protozoaea develops into zoaea. In some Anomura the egg hatches as a zoaea which passes through a metazoaea stage to become the adult. Metazoaea is, in fact, an advanced stage of zoaea but differs from it in having uniramous rudiments of thoracic appendages behind the maxillipede.
However, the third maxillipedes are biramous in hermit crab’s (Anomura) metazoaea and uniramous in that of crab (Brachyura). The abdominal appendages, i.e., pleopods also develop as buds. In some decapods, e.g., crabs, the life history starts from zoaea stage.
Larval Form - 5. Cypris Larva:
Cypris larva is covered by a bivalved shell having adductor muscle. Head has compound eyes, antennules with discs on which cement glands open, antennae are lost but remaining cephalic appendages are present, thorax has six pairs of biramous limbs, there is an abdomen of four segments. It has many adult features.
In Cirripedia, e.g., Lepas, the egg hatches as a nauplius, it changes into a cypris which gets fixed by discs of antennules with the secretion of cement glands, then it becomes a pupa which forms shell plates and rotates to assume the adult form.
Larval Form # 6. Mysis or Schizopod Larva:
Mysis or schizopod larva resembles an adult Mysis. Head and thorax have a carapace, all cephalic and thoracic appendages are present, but all thoracic appendages are alike and biramous with exopodites, abdomen has five pairs of pleopods and the sixth form uropods.
In some Decapoda, e.g., in Penaeus, a marine prawn, the egg hatches as nauplius, it passes by successive moults through zoaea stage, protozoaea stage and mysis stage which changes into an adult. In some lobsters, e.g., Homarus both nauplius and zoaea are passed within the egg, it hatches as a mysis larva which changes into an adult.
Larval Form ---7. Megalopa Larva:
Megalopa larva has a large un-segmented cephalothorax with all 13 pairs of appendages like those of a crab, abdomen is straight and in line with cephalothorax, it is like the abdomen of prawn with 6 pairs of well formed pleopods, In crabs the nauplius is passed in the egg, it hatches as a zoaea which by moulting forms the megalopa stage, the megalopa by moulting forms the adult.

In Decapoda there is a gradual abbreviation of development. Stages which are free larval forms in lower types of Crustacea are hurried through within the egg before hatching.
Larval Form--- 8. Phyllosoma Larva:
In Palinurus (the rock lobster), the egg hatches directly into a delicate, transparent, extremely flattened leaf-like larva called phyllosoma or glass crab. This larva is large sized having three distinct regions in the body, the head, thorax and abdomen. An oval carapace covers the head and a part of thorax. It possesses a pair of stalked compound eyes placed anterolaterally in the head.
Thorax bears six pairs of appendages; the first thoracic or maxillipedes are rudimentary, second are uniramous, third well formed biramous, and remaining three (4th, 5th, and 6th) pairs are biramous legs which are enlarged. Abdomen shows segmentation but appendages are absent. This larva undergoes several moultings and transforms into the adult. Phyllosoma is, however, considered to be modified mysis larva.

Larval Form # 9. Alima Larva:
In some Malacostraca like Squilla, the egg directly hatches out in a young stage called alima larva. It is a pelagic form having slender body with a short but broad carapace. Its body is glassy and transparent. In addition to all cephalic appendages, only first two thoracic appendages are found.
The abdomen has distinct six segments with four or five pairs of pleopods. The alima larva is supposed to be modified zoaea stage but it differs strikingly from zoaea in having the armature of the telson and well developed large second maxillipedes.
In addition to these, the glaucothoe larva of hermit crab resembles the megalopa larva as described earlier. Likewise, the calyptosis larva of Euphausia (a malacostracan) is similar to the zoaea larva in all essential features except that it possesses sessile paired eyes in place of stalked eyes.
Importance of Larval Stages:
The importance of larval stages may be accounted as under:
1. They help in wide dispersal of the species.
2. The larval stages help in establishing relationships between various groups.
3. Occurrence of nauplius stage in all crustaceans connects the different representatives of this class together. As referred to, the nauplius establishes relationship of some obscure animals like Sacculina where adult has lost the characters of the class and even the phylum.
In fact, it is the presence of nauplius stage in its life history that connects Sacculina to class Crustacea and further the presence of cypris stage relates it to subclass Cirripedia.
4. If Haeckel’s law of recapitulation (which states that every organism during its development, i.e., ontogeny, repeats its evolutionary history, i.e., phylogeny) is considered true then it can be said that the nauplius stage represents the ancestral form of crustaceans because all crustaceans invariably pass through nauplius stage during their development.
Hence, it can be concluded that present day crustaceans have evolved through nauplius stage.

1.2 Causes of Depletion of wildlife resources



a. Destruction of habitat: The natural habitat may be destroyed by man for his settlement, grazing grounds, agriculture, mining, industries, highway construction, drainage, dam building etc. as a consequence of this the species must either adapt to the changes, move elsewhere or may succumb to predation, starvation or disease and eventually die.
Construction of Dams: In India due to the construction of 1877 large and small scale dams all over india between 1980 and 2000 resulted in the submerging of 4.5 million hectors. No reforestation attempts were successful so far. The narmada vally project resulted in the submerging of 40000 hectors of forests in Madhyapradesh. A study done on the Indira Sagar Project of MP on the effects on wildlife showed that nearly 30% of the wildlife vanished due to project flooding.
Construction of dam on a river can block or delay upstream fish migrations and thus contribute to the decline or even the extcition of species. Construction of Hydroelectric Dam in Nepal cuts off local fish species eg. Tor and bangarius bagarius from their valuable spawining grounds upstream which resulted in decreasing of their numbers drastically.

b. Hunting for commercial products/trophies the wild animals are killed for their products such as hides and skin, tusks, fur meat, pharmaceuticals, perfumes, cosmetics and decoration purposes.
For example. In Africa, inrecent years 95% of the black rhino populations have exterminated by poachers for their horn. Today rhino hrn fetches upto 25000 dollars in pharmaceutical markets. In the last 10 years over one third of Africa’s elephants have been wiped out for some 3000 tonnes of ivory. In the international market the cost of ivory is 1000 dollars per kg. The major buyers are japan, honkong followed by US, Germany and UK. Indian government banned African Ivory in 1992
The scarlet macaw once common throughout south america has been eliminated from most of its range in central america. Several species of spotted cats as the ocelot and jaguar have been declining in numbers for the demand for their fur
In 1962, nearly 70000 whales were slaughtered. However international trade in whale products is banned now.
In india Rhino is hunted for its horns and musk deer for musk, elephant for ivory, Gharial and crocodile for their skin and jackal for fur. CITES lists nine indian animal species whiche have been depleted due to international trade. They are Fin Whale (Balenoptera physalus), Himalayan musk deer (Moschus moschiferus), Green turtle (Chelonia mydas), Desert Monitor Lizard (Varanus griseus), Yellow monitor lizar (Veranus flavescens) and Bengal monitor Lizard (Veranus bengalensis

c. Over Exploitation Excessive harvesting of marine organisms such as fishes, molluscs, sea cows and sea turtles has resulted in extinction of these animals
D. Collection for Zoo and research: animals and plants are collected throughout the world for zoos and biological laboratories for study and research in science and medicine. For example primates such as monkeys and chimpanzees are used for research as they have anatomical, genetic and physiological similarities to human beings.
E. Introduction of exotic species: Native species are subjected to competetion for food and space due to introduction of exotic species. For example, introduction of goats into galapogas islands in 20th century, resulted in destruction of habitats of several plants, birds and reptiles.
F. Control of pests and predators: predator and pest control measures, generally kill predators that are a component of balanced ecosystem and may also indiscriminately poison non target species.
G. Pollution/poisoning: Pollution alters the natural habitat. Water pollution especially injurious to the biotic components of estuary and coastal ecosystems. Toxic wastes entering the water bodies disturb the food chain and so that the aquatic ecosystems, insecticides and pesticides too affect adversely the plant and animal species.
h. hunting for food and fun: Hunting of wild animals takes place for the purpose of food because they form rich source of protein. Shikars are organized to kill wild animals for just fun and sport. Such activities decrease the number of wild life.
Other factors that contribute to the depletion of wild life are as follows
a. Distribution Range: the smaller the range of distribution the treater the threat of extinction
b. Degree of specialization: the more specialised an organism is the more vulnerable it is to extinction
c. Position of the organisms in the food chain: the higher the organisms is in the food chain, the more suceptible it becomes extict
d. Reproductive rate: large organisms tend to produce fewwer offsprings a widely spaced intervels

Thursday, August 24, 2017

What is the need of wildlife conservation?


a. Conservation preserves the ecological diversity and our life support systems water air and soil. If a species is lost, in long run, it may upset the natural balance and as a consequence makes the system vulnerable.

b. It also preserves the genetic diversity of plants and animals for better growth of species and breeding. All the native species possess genetic reservoir and provide genes for improvement of existing plant and animal life to solve present food problems or production of disease resistant varieties.
c. In agriculture, we are still dependent on traditional crop varieties. Fisheries too are heavily dependent on the maintenance of aquatic biodiversity
d. AFTERALL THIS EARTH BELONGS TO NOT ONLY FOR HUMANS BUT ALSO FOR ALL ORGANISMS.
e. Economic benefit: wilelife is a source of income to recreation and tourism industry. Most popular tourist atractions are the wildlife sanctuaries and national parks.
Many plants have medicinal value. Some animals like shrimps and crabs provide medicines against fungal infections
f. Game value. Wildlife has its worth as game also. In several western countries, million of people hunt or fish for recreation, spending millions of dollars for such sport
g. Scientific value: scientists of wildlife in biology and medicine are direct value for humans. For examples sea urchins have helped greatly in the understanding of human embryology, rhesus monkeys in present blod groups, antlers of deer in determining the degree of radio active contamination of environment.
h. Aesthetic value: aesthetic values such as the taste of wild berries, fragrance of wild flowers compel us to preserve them.
A world without melodious birds, graceful beasts and thick forest would be poorer place for humans to live in.

Thursday, August 3, 2017

PARENTAL CARE

PARENTAL CARE IN AMPHIBIA
Amphibian include anurans, urodelans and apodans. In all these groups of amphibians we come across with a great deal of parental care. Amphibians show several mechanisms to protect their eggs and developing young ones because of the they lay few eggs.










Parents protect the eggs and early developmental stages in two ways.
1) They construct nests
2) Direct Nursing.
The female Icthyophis glulinosa will dig a hole in the moist soil near a pond. It will deposit eggs in it. Around this egg mass the mother will coil and. protect the egg mass from the enemies.

3. PARENTAL CARE IN ANURA AMPHIBIANS:

In Mura amphibians the parental care is reached its peak. Many organisms will exhibit parental care.
1) Protection by Nests:
Many frogs and toads build nests in which the eggs are laid and developed. This is a primitive method of parental care. In these organisms the larva comes out in a very early embryonic stage which requires some kind of protection in the very early stages of development, hence the parent will build nests.
I) Hyla Faber:- It is Brazilian free frog. The female will construct the nest in the shallow waters of a pond. The female will dig a hole of 8 to 10cm depth. The mud which comes out of it is used by the female Hyla to construct a wall around the hole. This wall is raisd’above the level of water. Female Hyla will make the inner surface of this Nursery smooth and even the female will lay eggs in this nursery. The eggs and larval forms are protected inside this structure.
ii) Rhacophorus malabaricus It is called chunam frog. It lays eggs on the branches or leaves of a tree which will be hanging over a pond. These larvae after hatching from eggs will fall into the pond water and undergo metamorphosis.











lii Rhacophorus schlegeli It is called Japan free frog. Both male and female frog will make a burrow in the moist soil near a pond edge .This hole is filled with foam by female. Then female lays eggs in this foam. The male and female animals will make an exit tunnel into the pond from the hole. The larvae developed from eggs will be carried by the liquid formed from the foam into the pond through these exit tunnels.

lv) Hyla resinfectris It is a free frog. It will make use of holes in the frees. It will line the hole of free trunk with beewax brought from bee comb. Female animal lays eggs in this hole when filled with rain water.

v) Leptodactylus mystacinus It digs a. hale in the moist mud near a pond. This hole is filled with foam which is secrete4 by the oviducts of the female. Female lays eggs in it. These eggs hatch into larvae then the pond will get good amount of water from rain. Then these holes are also sunounded by the pond water, Then the larvae will enter the pond and grow.
vi) Hyla nebulosa It lays its eggs in a nest. This nest is made by dry leaves. The eggs hatch and develop into small adults. Larval form is absent.
vii) Hylodes : It is an American frog. It deposits its eggs in moist places or under the stones. The eggs hatch and give tiny adults in perfect condition.

2) Direct Nursing by Parents:
This method is more advanced than protecting the eggs and larvae by constructing nests.
1) Alytes . It Is called Mid-wife-toad When the eggs are laid by the female frog winds the strings of eggs round his back and thighs. This male frog lives in a shallow pit of the moist soil, It will came out of the pit now and then for feeding and to make the eggs moist. When the eggs are ready to hatch the male frog moves to a near by pond and the larvae are released.







2. Gastrotheca It is called New World brooding (or) Marsupial frog. It has a special pouch in its skin. It opens out through an opening near the cloaca. Fertilized eggs are transferred into this pouch. The eggs are stored in this pouch where they ‘undergo development and tadpoles are liberated out.
3, Pipa americana It is called surinam toad. During breeding season the skin of the back of the female becomes soft and spongy. During copulation the oviducts will come out of the female- Because of the movements of male the eggs are forced out of the oviduct. Each egg sinks into a small pocket of the skin. It gets coveted by an operculum. In the soft maternal tissue the young one can develop safely. The developing embryo has a tail and yolk sac. It has no gills. The tail may work like placental connection to draw nutrition from the mother. Nearly after 80 days small individuals may come out.
4. Rhinoderma darwini It is called little South American frog. The fertilized eggs are transferred into its vocal sacs the development takes place. Then the completely developed young individual will jump out from the mouth of the male frog.
Thus in Amphibians a good amount of parental care is seen.


Saturday, July 29, 2017

NERVOUS SYSTEM OF PILA

NERVOUS SYSTEM OF PILA

Nervous system of Pila consists of paired ganglia, commissures and connectives uniting them, and nerves running from these central organs to all parts of the body.

1. Ganglia: Paired ganglia which are groups of nerve cells are as follows.

a) Cerebral ganglia: A pair of roughly triangular ganglia, situated anteriorly on the dorso-lateral sides of the buccal mass.

b) Buccal ganglia: A pair of small, triangular ganglia, lying dorso-laterally one on either side at the junction of the buccal mass and oesophagus, partly embedded in the muscles.

c) Pleuro-pedal ganglia: A pair of large, somewhat triangular ganglionic masses present one on either ventro-lateral side of the buccal mass.

 Each one is formed by the fusion of an outer pleural and an inner pedal ganglion, separated by a faint notch.

 Right pleuro-pedal ganglionic mass also consists of the infra-intestinal ganglion fused with it.

d) Supra-intestinal ganglion: An unpaired fusiform ganglion, lying in a sinus behind the left pleuro-pedal ganglionic mass.

e) Visceral ganglia: A single ganglionic mass representing two fused ganglia, situated at the lower end of the visceral mass.

2. Commissures: Commissures are those nerves which establish connections between two similar ganglia and lying dorsally to the buccal mass.

b) Buccal commissure: A fine nerve which connects the two buccal ganglia and runs transversely on the ventral side of the oesophagus.

c) Pedal commissures: Two thick nervous bands that lie one above the other underneath the buccal mass and connect the two pedal ganglia together.

3. Connectives: Connectives are those nerves which connect two different ganglia. In the nervous system of Pila, these are

a) Two cerebro-buccal connectives: These connect, on either side, the cerebral ganglion and buccal ganglion together.

b) Two cerebro-pleural connectives: These connect, on either side, the cerebral and outer pleural ganglion of the pleuro-pedal ganglionic mass.

c) Two cerebro-pedal connectives: These connect, on either side, the cerebral and inner pedal ganglion of the pleuro-pedal ganglionic mass.

d) Pleuro-infra intestinal connective: Also called infra-intestinal nerve, it is a nerve connection between the pleural ganglion of the left pleuro-pedal mass and the infra-intestinal ganglion which is fused with the right pleuro-pedal mass.

e) Infra-intestinal visceral connective: Running below the intestine, it is a long nerve that connects the visceral ganglion with the infra-intestinal part of the right pleuro-pedal-infra-intestinal ganglionic mass.

f) Supra-intestinal visceral connective: Running above the intestine, it is a slender nerve that connects the visceral ganglion with the supra-intestinal ganglion.

g) Supra-intestinal-pleural connective: Also called the supra-intestinal nerve, it connects the supra-intestinal ganglion with the pleural part of the right pleuro-pedal-infra-intestinal ganglionic mass.

h) Zygoneury: It is a nerve connection between the pleural part of the left pleuropedal ganglionic mass and supra-intestinal ganglion.

4. Nerves to different parts of the body: Various ganglia send nerves to innervate different parts of the body

i) Each cerebral ganglion gives off nerves, supplying the snout, skin, tentacle and buccal mass anteriorly and the tentacle, eye and statocyst, posteriorly.

ii) Buccal ganglion of each side , sends nerves to innervate the buccal mass, radular sac, salivary glands, oesophagus and oesophageal pouches
iii) Pedal ganglia gives off nerves, anteriorly as well as posteriorly, to innervate the foot. Statocyst, on each side, is also connected, by a band of, connective tissue, to each pedal ganglion.

iv) Left pleural ganglion innervates the parietal wall, mantle, osphradium, left nuchal lobe, columellar muscle and anterior part of the gill.

v) Pight pleural ganglion innervates the parietal wall, epitaenia, right nuchal lobe, copulatory organ, columellar muscle and rectum.

vi) Supra-intestinal ganglion gives off a stout nerve to innervate the mantle and the anterior part of ctenidium, while its connective with the left pleural ganglion sends a few nerves to the parietal wall.

vii) Nerves from the visceral ganglion supply the renal organ, genital organs, pericardium, stomach, intestine, digestive gland etc.

Sense organs

A snail is diffusely sensitive for the sensory cells are distributed all over the head, foot and various other parts of the body. Special sense organs of sense include a single osphradium and paired eyes, statocysts, labial palps and tentacles.

I) Osphradium: Single osphradium is situated on the left side of the animal suspended from the roof of the mantle cavity close to the entrance through the left nuchal lobe.

 It is a small, elongated, oval structure, about 6-7 mm long. While broadest in the middle, its inner or right end is bluntly rounded and the outer or left end is somewhat pointed.

 It is bipectinate consisting of 22-28 thick, fleshy, and roughly triangular leaflets, arranged in two rows along a slightly raised median or central axis.

 Leaflets are largest in the middle of the osphradium. Each leaflet is attached to the mantle wall by its broad base, to the central axis by its smaller inner side, while its outer longer side remains free.

 Osphradium is supplied by a nerve from the left pleural ganglion.


 In a transverse section, the osphradium consists of an outermost covering of a single layered epithelium, internally lined by a thin basement membrane, the interior filled up with nerves, connective tissue and blood spaces.

 Epithelial cells are elongated possessing basal nuclei, and they are of three types
i) sensory
ii) ciliated
iii) glandular

 The ciliated cells line the attached margin, while the sensory cells are with out cilia, cover the osphradium. Flask-shaped glandular cells are found scattered among the sensory cells.

 Osphradium hangs like a curtain in the path of the respiratory water current, and probably serves as an olfactory organ. Its name has been derived from a Greek word, meaning to smell.

 It serves to test the chemical nature of the inspiratory water current. In case the water is foul, its entry into the mantle cavity is stopped by the closure of the left nuchal lobe.

 It may also help in the selection of the food material.

II) Eyes:
 Snail’s head carries a pair of short fleshy and stump-like stalks or ommatophores, one on either side, behind the second pair of tentacles.

 Each ommatophore bears a small, black and circular eye, slightly below its tip on the outer side.

 In spite of their elaborate structure, eyes of Pila are probably not true organs of sight. Sense of sight is greatly limited in range and the snail does not seem to distinguish objects, but only responds to changes in the intensities of light and detect quick movements.

 Most of the snails feed at night probably because their eyes are adapted to dim light. In some snails the lost eyes can be replaced by regeneration, a process which has been recorded to occur 20 times in succession.

III) Statocysts:
 Statocysts are a pair of small, pyriform and cream-coloured sacs, lying one on either side attached to the pedal ganglion of that side by a band of connective tissue.

 Each statocyst lies in a depression, posterior and outer to the ganglion. Each statocyst is a hollow capsule surrounded by an outer thick, tough, leathery covering of connective tissue.

 Wall of the capsule is made of a single layer of ectodermal cells, and supplied by a nerve from the cerebral ganglion.

 Cavity of the capsule is filled with a fluid and a variable number of minute, oval and calcareous particles the statoconia. Statocysts are organs of equilibrium.

IV) Tentacles
 The snout of Pila is anteriorly prolonged into a pair of short, contractile and conical processes bordering the mouth.

 These are the labial palps or the anterior or first pair of tentacles.

 Behind them arise, one on either side, a pair of long, tapering filamentous and highly contractile whip-like processes, the true or second pair of tentacles.
 Tentacles are of the same colour as the snout and tactile in function. A sense of taste is doubtfully attributed to the labial palps.

Thursday, July 13, 2017

PRACTICAL NOTES

1. Eleutheronema tetradactylum
Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Polynemidae

Genus: Eleutheronema

Species: tetradactylum
1. Eleutheronema tetradactylum, also known as Indian Salmon or Rawas, is a threadfin fish of the Polynemidae family.
2. This highly commercial fish known for uses in aquaculture occur mainly over shallow muddy bottoms in coastal waters forming loose schools.
3. Adults of this highly vulnerable species enter rivers during winter.
4. Adults feed on prawns and fish with occasional polychaetes, while juveniles feed on prawns shrimps and mysids.

2. Epinephelus malabaricus
Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Serranidae

Genus: Epinephelus

Species: malabaricus
1. The Malabar grouper is widespread throughout the tropical waters of the Indo-West Pacific area from the eastern coast of Africa to the Tonga Islands, Red Sea included.
2. This grouper lives in various habitats, such as lagoons, mangroves, coral and rocky reefs, sandy and muddy bottom areas, between 2 and 150 m deep.
3. The juveniles prefers lagoon or brackish areas.
4. It has a light grey to light brownish background color, with a number of dark brown spots randomly scattered.
5. The body has also a various number of brown diagonal stripes, but in maturity they seem to become a uniform darker colour.
6. Young fish have numerous brown spots. The tail fin is rounded.




Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Clupeiformes

Family: Clupeidae

Subfamily: Alosinae

Genus: Hilsa

Species: ilisha
3. Hilsa ilisha

1. It is found in rivers and estuaries in India, Pakistan, Bangladesh, Burma and the
2. It has no dorsal spines but 18 - 21 Dorsal soft rays and anal soft rays. The belly has 30 to 33 scutes.
3. There is a distinct median notch in upper jaw.
4. Gill rakers fine and numerous, about 100 to 250 on lower part of arch and the fins are hyaline.
5. The fish shows a dark blotch behind gill opening, followed by a series of small spots along the flank in juveniles.
4. The species filter feeds on plankton and by grubbing muddy bottoms.

4. Labeo calbasu
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Cypriniformes
Family: Cyprinidae
Genus: Labeo
Species: calbasu
1. Labeo is a genus of carps in the family Cyprinidae. They are found mainly in the Old World tropics.
2. It contains the typical labeos in the subfamily Labeoninae, which may not be a valid group, however, and is often included in the Cyprininae as tribe Labeonini.
3. The labeos appear fairly similar to the "freshwater sharks" of the genus Epalzeorhynchos, which is also part of the Labeoninae (or Labeonini), but is not very closely related.
4. Labeos are larger, and have a more spindle-shaped body, as they are mostly free-swimming rather than benthic like Epalzeorhynchos.
5. Their mouths look very different, too; they have a pronounced rostral cap, which covers the upper lip except when feeding.
6. The lips are expanded into thick, sausage-shaped pads which have keratinized edges.


5.Megalops cyprinoides
Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Elopiformes

Family: Megalopidae

Genus: Megalops

Species: cyprinoides
1. The Indo-Pacific tarpon, Megalops cyprinoides, also known as the Oxeye herring or simply herring, is a relatively medium-sized species of tarpon.
2. In appearance, it is like the Atlantic tarpon, Megalops atlanticus: olive-green on top, and silver on the sides.
3. The large mouth is turned upwards; the lower jaw contains an elongated, bony plate.

4. The last ray of the dorsal fin is much longer than the others, reaching nearly to the tail.
5. It is capable of filling its swim bladder with air and absorbing oxygen from it.
6. Species in fresh water tend to be smaller than the saltwater species, growing just over 50 centimetres (20 in), while saltwater species grow over a 1 metre (3.3 ft).
6. They are an opportunistic feeder, feeding on smaller fish, crustaceans, and evenplants rarely.
Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Carangidae

Genus: Parastromateus

Species: niger
6.Parastromateus niger
1. The black pomfret, Parastromateus niger, is a species of carangid native to reefs of the Indian Ocean and the western Pacific Ocean.
2. It is found at depths from 15 to 105 m (49 to 344 ft), though it is rarely found deeper than 40 m (130 ft).
3. This species grows to 75 cm (30 in) in total length and is very important to local commercial fisheries. This species is the only known member of its genus.





Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Scombridae

Genus: Scomberomorus

Species: commerson
7. Scomberomorus commerson
1. They are vivid blue to dark grey in colour along their backs and flanks and fade to a silvery blue-grey on the belly.
2. Spanish mackerel have scores of narrow, vertical lines down their sides.
3. Spanish mackerel are the largest of all Australian mackerels, growing to about 200 cm and up to 70 kg.
4. Spanish mackerel spawn in oceanic conditions on reef edges.
5. Eggs have a large oil droplet that aids in buoyancy and keeps them at the top of the water column which is warmer, well oxygenated, and has an abundant planktonic food supply for the larvae once they are hatched.
8. Stromateus argenteus
Kingdom: Animalia

Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Stromateidae

Genus: Pampus

Species: argenteus
1. Pampus argenteus, often called either the silver or white pomfret, is a species of butterfish that lives in coastal waters off the Middle East, South Asia, and Southeast Asia.
2. Fish of this family are characterized by their flat bodies, forked tail fins, and long pectoral fins.
3. Silver pomfrets are usually silver/white in color, with few small scales.
4. They can grow up to 4-6 kg. However, due to overfishing, specimens weighing less than 1 kg are more commonly seen.
5. It is called pamplet in Mumbai.










Tilapia mossambica
Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Cichlidae

Genus: Tilapia

Species: mossambicus
1. The Mozambique tilapia, Oreochromis mossambicus, is a tilapiine cichlid fish native to southern Africa.
2. It is a popular fish for aquaculture. Dull colored, the Mozambique tilapia often lives up to a decade in its native habitats.
3. This makes it an optimal species for aquaculture because it readily adapts to new situations. It is known as Black Tilapia in Colombia and as Blue Kurper in South Africa.
4. The native Mozambique tilapia is laterally compressed, and has a deep body with long dorsal fins, the front part of which have spines.
5. It is a remarkably robust and fecund fish, readily adapting to available food sources and breeding under suboptimal conditions.
Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Rachycentridae
Genus: Rachycentron
Kaup, 1826

Species: R. canadum
Rachycentron canadum
1. The cobia (Rachycentron canadum) is a species of perciform marine fish, the only representative of the genus Rachycentronand the family Rachycentridae.
2. The another common names include black kingfish, black salmon, ling, lemonfish, crabeater,prodigal son and aruan tasek.
3. Attaining a maximum length of 2 m (78 in) and maximum weight of 78 kg (172 lb), the cobia has an elongated fusiform (spindle-shaped) body and a broad, flattened head.
4. The eyes are small and the lower jaw projects slightly past the upper. Fibrous villiform teeth line the jaws, the tongue, and the roof of the mouth.
5. The body of the fish is smooth with small scales. It is dark brown in color, grading to white on the belly with two darker brown horizontal bands on the flanks.
Scomberomorus guttatus
Phylum: Chordata

Class: Actinopterygii

Order: Perciformes

Family: Scombridae

Genus: Scomberomorus

Species: guttatus

1. Indo-Pacific king mackerel or popularly (spotted) seer fish (Scomberomorus guttatus) is a sea fish among the mackerelvariety of fishes. It is found in around the Indian ocean and adjoining seas.
2. It is a popular game fish and grows up to 45 kg (100 lbs) and is a strong fighter, that has on occasion been seen to leap out of the water when hooked.
3. It is very popular among the countries of the Indian subcontinent including India, Sri Lanka and Bangladesh. It's a fairly expensive fish that's considered a delicacy in most places.


Monday, July 10, 2017

POLYMORPHISM IN COELENTERATES

POLYMORPHISM IN COELENTERATES
 The presence of polymorphism in cnidarians is one of their characteristic features. It is defined as the occurrence of structurally and functionally different types of individuals within the same organism during its life cycle.
 A species that exhibits this phenomenon is called polymorphic.
 Polymorphism is predominantly exhibited by the different animals of class- hydrozoa.
 Hydroid colonies which bear two types of zooids are known as dimorphic, while colonies which bear more than two types of zooids are called polymorphic colonies.
Two basic forms
 In Hydrozoa (or Coelenterates), which may be single or colonial, there occur two main types of individuals or zooids- polyps and medusae.
 1. Polyp: It is sessile (fixed to the substratum) with a hydra like body attached to the main colony by narrower end. Its free end is wider and raised into hypostome that bears a mouth surrounded by tentacles. It faces upwards and carries the function of feeding the colony. Polyps are specialized for feeding and thus known as gastrozooids.






 2. Medusa: It is a free swimming zooid with an umbrella shaped body having exumbrellar and subumbrellar surfaces. It has a mouth facing downwards (present on the tubular growth called manubrium hanging down from subumbrellar surface) in contrast to polyp in which mouth faces upwards.
 Like hydranth, it can feed on its own for its survival until its function is over. It bears four gonads (testis or ovary) which produces either sperms or ova at the time of maturity and is responsible for sexual reproduction therefore also known as gonozooid. They normally die after reproducing the colony. Therefore, they not only help in sexual reproduction but also play an important role in dispersal of the colony.



Importance of Polymorphism
 Polymorphism is essentially a phenomenon of division of labour. Different functions are assigned to different individuals, rather than to parts or organs of one individual.
 Thus, polyps are concerned with feeding, protection and asexual reproduction, while medusae are concerned with sexual reproduction.
Patterns of polymorphism

Degree of polymorphism varies greatly in different groups of Hydrozoa
1. Dimorphic. Simplest and commonest pattern of polymorphism is exhibited by many hydrozoan colonies like Obelia, Tubularia, Campanularia etc.
 They have only two types of zooids. Gastrozoids or hydranths are concerned with feeding. While gonozoids or blastostyles with asexual budding forming sexual medusae or gonophores.
 Such colonies, bearing only two types of individuals are called dimorphic and the phenomenon is termed dimorphism.
2. Trimorphic. Some forms like Plumularia, are trimorphic. Besides Gastrozooids and gonozooids, they also possess a third type of individuals, the dactylozooids. These are functionally non-feeding and defensive polyps bearing batteries of nematocysts.
3. Polymorphic. Coelenterates having more than three types of individuals are called polymorphic.
 A somewhat greater degree of polymorphism is found in the encrusting colony of Hydractinia with five types of polyps each performing a specialized function.
These are: 1. Gastrozooids for feeding
2. Spiral dactylozooids for protection
3. long sensory tentaculozooids with sensory cells
4. skeletozooids as spiny projections of chitin
5. gonozooids or reproductive individuals, bearing male or female gonophores or medusae for sexual reproduction




Extreme examples of polymorphism are seen in the pelagic or swimming colonies of the orders
 Siphonophora (Diphes, Halistemmia, Stephalia, physalia) and Chondrophora (porpita, velella).
 Polymorphis reaches its peak in siphonophora.









(a) Modifications of polyps
Polyps structurally get modified into different types of zooids according to the requirement of an individual, which are described below:
i. Gastrozooids: These are feeding zooids and resemble the structure of polyp without usual tentacles. They are tubular, elongated, with a mouth facing towards the bottom of the colony.
ii. Dactylozooids: These are protective zooids. They may be also called as feelers or palpons. Structurally, they look like gastrozooids but are blind structures without any mouth.
iii. Gonozooids: They may resemble gastrozooids having mouth but are without tentacles and bear medusa. In other the gonozooids may form stalked branches bearing grape like structures called gonophores. Sometimes tentacles like dactylozooids are attached to them which are called gonopalpons.
iv.Pneumatophore:
It is a hydrostatic apparatus present in siphonophores. It is gas filled chamber that appear to be a highly modified polyps (although previously considered as derived from medusae).
 It helps in keeping the body in an upright condition while floating.
 It is without mesogloea but the umbrella cavity contains an air chamber called a pneumatocyst (Fig. 8a). Cells lining the pneumatocyst secrete the gases or may expel out of it through one or more small openings called stigmata.
 Thus pneumatophore is a balloon like structure or a hydrostatic chamber containing air.
 When pneumatophore is filled with air, the colony becomes lighter and floats at the surface of the water, but when the gas is expelled out of the pneumatophore, colony sinks down.






b. Modifications of medusae . The medusoid individuals are of the following types
1. Nectophore or nectocalyx or swimming zooid with a muscular bell without manubrium or tentacles.
2. Pneumatophore or float as a bladder-like medusa filled with secreted gas.
3. Phyllozooid or bract, usually leaf-like and studded with nematocysts, serving for protection of other zooids.
4. Gonophore bearing gonads, which may be either male, producing sperms, or female producing ova.









Origin of polymorphism
 As we have seen, colonies of Siphonophora represent the most specialized of Hydrozoa attaining the highest degree of polymorphism and presenting the greatest number of medusoid and polyploid types.
 There are two views regarding which came first, polyp or medusa, during the evolution of polymorphism in Coelenterata.
 According to one view, the ancestral coelenterate was a hydra-like polyp which arose from gastraea. It gave rise to hydroid colony by asexual budding. In the sessile colony some polyps became modified into medusae for sexual reproduction and pelagic life.
 Thus, through division of labour, the hydroid colony became polymorphic.
 According to second view (Brooks, 1886), which seems to be more acceptable, the ancestral coelenterata was a primitive medusa.
 It arose from metagastreae by developing tentacles and becoming free swimming.
 According to Huxley, and Metschnikoff, the manubrium, tentacles and umbrella of this p0rimitive medusoid individual were multiplied and shifted from their original positions to become various zooids of the polymorphic colony.
 According to this view, polyploid stage is considered the persistent larval stage and medusoid the completely evolved coelenterate.

















CORALS AND CORAL REEF FORMATION
 Meaning of corals: Coral animals or corals are marine, mostly colonial, polyploid coelenterates, looking like miniature sea anemones and living in a secreted skeleton of their own.
 Their calcareous or horny skeleton is also commonly known as coral.
 Some corals grow into massive, solid structures others form large, branched colonies.
 Most of the corals belongs to the class Anthozoa and a few to the class Hydrozoa of phylum coelenterata.
Structure of coral polyp:
1. soft structure
 A typical coral polyp from a colony is a small organism about 10mm long and 1to 3 mm in diameter. Solitary coral polyps are much larger reaching up to 25 cm in diameter.
 A basal disc is absent because the basal region of polyp is surrounded by calcareous exoskeleton.
 Oral disc bears numerous tentacles, in several rows around an elongated, oval or circular mouth. Pharynx or stomodaeum is short and without siphonoglyphs.
 Mesenteries are restricted to the upper part of coelenteron and mesenterial filaments contain only one glandular lobe bearing nematocysts.
 A typical coral polyp from a colony is a small organism about 10mm long and 1to 3 mm in diameter. Solitary coral polyps are much larger reaching up to 25 cm in diameter.
 A basal disc is absent because the basal region of polyp is surrounded by calcareous exoskeleton.
 Oral disc bears numerous tentacles, in several rows around an elongated, oval or circular mouth. Pharynx or stomodaeum is short and without siphonoglyphs.
 Mesenteries are restricted to the upper part of coelenteron and mesenterial filaments contain only one glandular lobe bearing nematocysts.







 Living polyps are found only on surface layers of coral masses. They feed at night both by raptorial and suspension feeding. When not feeding they withdraw into cup-like cavities of skeleton.
Structure of coral skeleton:
 Skeleton of a solitary coral is known as corallite. It is a calcareous exoskeleton secreted by epidermis.
 In a colonial coral corallites of individual polyps fuse together to form a skeletal mass, called corallum.
 Each corallite is like a stony cup with a basal part or basal plate, and a cup wall or theca, enclosing the aboral portion of the polyp.
 Cavity of cup contain a number of vertical radiating ridges called sclerosepta, proceeding from theca towards the centre of the cup.








 Inner ends of sclerosepta are fused to form an irregular central skeletal mass or columella.

Formation of coral skeleton:
 In coral polyps sexual reproduction takes place by fusion of gametes. Zygote develops into a free swimming ciliated planula larva which settles down and metamorphosis into a young coral polyp.
 There is no medusa stage. By asexual budding, single become s the parent of all other members of the colony.
 The coral polyp begins to secrete a skeleton rudiment or prototheca. It is secreted by ectoderm, first as a basal plate. Following it, radial folds develop which secrete sclerosepta. At the same time a rim is built up as a thecal wall around the polyp, lying at the top.
 Mean while further skeletol material is added into the gaps between sclerosepta of skeleton which usually alternates with mesenteries of the polyp.
 Coral colony grows in size continuously by budding of new polyps, particularly along the margins and on surface layers of coral masses. Variety in form of compound corals results due to cavities patterns of budding.


Coral reefs
Meaning of coral reefs:
 Coral colonies grow continuously in size by budding of polyps and often forms extensive masses, known as coral reefs.
 A coral reef is a ridge or mound of limestone, the upper surface of which near the surface of sea and which is formed chiefly of CaCo3 secreted by coral polyps.
 Principal builders of coral reefs are stony corals (Madreporaria), but other important contributors are the hydrocorallines and alcyonarians. Coralline algae and Foraminiferan Protozoa also take part in the formation of coral reefs.










 Reef building corals require warm shallow waters (normally above 200C). They are therefore limited to the Indo-Pacific, the Central Western Pacific, and the Caribbean regions north of Bermuda.
Kinds of coral reefs:
There are three types of coral reefs:
• a. Fringing reefs
• b. Barrier reefs
• c. Atoll
 a. Fringing reefs: Fringing reefs are developed in shallow waters on or near the shores of the volcanic islands. These are the simplest kind of reefs. They extend from the sea shore towards the sea as a platform ranging from few meters to half a kilometer and then slopes down towards the bottom of the sea








 Fringing reefs consist of several zones that are characterized by their depth, the structure of the reef, and its plant and animal communities.
Reef edge or reef front
Seaward slope
Reef flat
Boulder zone
Lagoon
b. Barrier reefs.
 A volcanic island with the fringing reefs is surrounded with a big channel of water called lagoon. Its depth may vary from 20 to 100 meters and even ships can pass through it. Lagoon is further surrounded by reefs called barrier reefs.
 As their name suggests, they act as a barrier for ships between sea shore and the main sea.
 Lagoon has fringing reefs towards the volcanic coast and barrier reefs on the other side of it. Sometimes both fringing reefs and barrier reefs may join each other at the bottom of the sea.
 There is a Great Barrier Reefs of Australia. It is not a single structure but is made up of many strings of separate reefs joined to each other at the bottom and thus forms a very big structure which extends along the north eastern coast of Australia for over 2000 kilometers.
 It is separated from the main land by a lagoon which is around 15 to 250 kilometers wide and 70 meters deep. During high tide, big ships can sail over it without realizing that reefs are present nearby and may crash. The Great Barrier Reef is the contribution of all different kinds of coral growth over the years.








 c. Atoll: Atolls are coral reefs which are present within sea water hundreds or thousands of kilometers away from the nearest sea shore.
 There is no volcanic island present. It is more or less circular or horse shoe shaped reef enclosing a central lagoon which may be 40 or 50 miles across and 20 to 90 meters deep.
 It may be a complete or broken into many small reef islands separated from each other by water channels.
 At some places reef is low so sea water simply covers it and reef is not visible. Sometimes a large atoll is formed by many small islets joined together along a line of reef. Thousands of such atolls are found in the South Pacific.
 It must be noted that reefs are not continuous rigid structures but they are broken up into many reefs and islands by water channels. Suvadiva is the largest atoll present in Maldives.
 Bikini Atoll has 2.87square miles land area with a lagoon area of 280 square miles. It was inhabited by the people but people moved to different places as it was selected by the United States for testing hydrogen and atomic bombs. Horse shoe shaped atoll of West Texas is 70 to 90 miles across and 1,000 meters thick.









Theories explaining the formation of coral reefs:
a. Darwin’s subsidence theory: Darwin believed that the reef began as fringing reefs on a sloping shore. Then the island subsides very slowly, so slowly that the reef grows upward at about the same rate, naturally the island becomes small, the channel between the reef and land widens and thus the fringing reef transforms into a barrier reef. Further subsidence of the land till it sinks completely out of site results in the formation of an atoll (Fig. 18). This is substantiated by the fact that all the known coral reefs were in regions where a sinking of the land was known to have taken place or where there were evidences that it had probably occurred.
b. Samper Murray solution theory: This theory states that calcareous skeletons of animals and other sediments form big mounds on the floor of the oceans. Over the time, these deposits grow to certain heights and corals grow on them and reach the water surface. Barrier reef is formed by the luxurious growth of coral at the outer edge while atoll is formed by dissolution of the inner coral rock.
c. Submerged bank theory This theory states that both barrier reefs and atolls grew upon pre-existing flat surfaces. Extensive coral growth occurred on a flat surface which got submerged in the water with the passage of time. Exposed regions formed the barrier reefs, while the shape of the atolls is obtained by the action of prevailing water currents and winds.













Significance of corals:
i. They show a high degree of physiological integration, division of labor, and perfect coordination with other groups of animals staying together, mutually benefiting each other in getting shelter, protection and food. Thus they constitute an ecologically important aquatic ecosystem.
ii. They help in studying the evolution of the animals as fossils of the animals are preserved in the coral reefs over the years.
iii. By studying the lines of growth on the fossils of some mollusk help in knowing about the seasonal fluctuations as the thickness of the growth line varies from season to season.
iv. A few stony corals because of the presence of minute pores are used in surgical procedures as human capillaries can easily pierce through equal sized pores which are helpful in interconnecting two bones with each other. These corals are being used in the surgery for bone grafts and jaw surgery etc.
v. Horseshoe shaped Atoll is the largest limestone reservoir for the oil production in North America.
vi. Many of the corals are precious stones which are used in making jewelry and have aesthetic value.
vii. Coral reefs are very hard structures and are an important source of mortar, cement, lime etc as they contain enough amount of CaCO3, therefore their rocks can be used for making roads and houses etc.
viii. Reefs are also a rich source for medical formulations, used to treat a wide range of diseases like asthma, heart diseases, and viral, fungal and bacterial infections. It has been reported in 2006 that Yellow coral (Isis hippuris) collected off the coast of Okinawa island of Japan has yielded a compound that can slow down and possibly prevent virus replication and also treat cancer.
ix. Lastly, corals act as affective buffers against erosion and storms occurring in the sea thus help in preventing tsunami disaster.